Publication Archive


Gonzalez-Santoyo, I. Wheatley, J.R. Welling, L.L.M. , Cárdenas, R.A., Jimenez-Trejo, F., Dawood, K., and  Puts, D.A.  (2015). The face of female dominance: Women with dominant faces have lower cortisol.  Hormones and behavior , 71: 16-21. PDF

Puts, D.A. (2015). Human sexual selection.  Current Opinion in Psychology, 7: 28-32. PDF

Puts, D.A.
, Bailey, D.H., and Reno, P. (2015). Contest competition in men. In: The Handbook of Evolutionary Psychology, 2nd ed. Buss, D.M. ed. Wiley and Sons, pp.385-402. PDF

Puts, D.A.
Pope, L.E.Hill, A.K., Cárdenas, R.A.,  Welling, L.L.M.Wheatley, J.R., and Breedlove, S.M. (2015). Fulfilling desire: Evidence for negative feedback between men’s testosterone, sociosexual psychology, and sexual partner number.  Hormones and Behavior, 70: 14-21. PDF

Wheatley, J.R.
 and  Puts, D.A. (2015). Evolutionary science of female orgasm. In:  The Evolution of Sexuality, Shackelford, T.K.and Hansen, R.D. (eds.). Springer, pp.123-148. PDF


Bossio, J., Suschinsky, K.,   Puts, D.A. , and Chivers, M. (2014). Does menstrual cycle phase influence the gender specificity of heterosexual women's genital and subjective sexual arousal?   Archives of Sexual Behavior , 43(5): 941-952.  PDF

Claes, L.P., Liberton, D., Daniels, K., Matthes Rosana, K., Quillen, E., Pearson, L., McEvoy, B., Bauchet, M., Zaidi, A.A., Yao, W., Tang, H., Barsh, G., Absher, D.M.,   Puts, D.A. , Rocha, J., Beleza, S., Pereira, R.W., Baynam, G., Suetens, P., Vandermeulen, D., Wagner, J.K., Boster, J., and Shriver, M.D.; (2014). Modeling 3D facial shape from DNA.   PLoS Genetics . 10(3):e1004224.  PDF

Doll, L.M.,  Hill, A.K.
, Cárdenas, R.A.,   Welling, L.L.M. Wheatley, J.R. , and   Puts, D.A.  (2014). How well do men’s faces and voices index mate quality and dominance?   Human Nature , 25: 200-212.  PDF

Hodges-Simeon, C.R., Gurven, M.,  
Puts, D.A. , and Gaulin, S.J.C. (2014). Vocal fundamental and formant frequencies are honest signals of threat potential in adolescent males.   Behavioral Ecology , 25(4): 984-988. PDF

Li, Y., Bailey, D.H., Winegard, B.,  
Puts, D.A. ,   Welling, L.L.M. , and Geary, D.C. (2014). Women’s preference for masculine traits is disrupted by images of male-on-female aggression.   PLoS ONE , 9(10): e110497.  PDF

Puts, D.A.
 (2014). Review of   Evolution and Human Sexual Behavior , Gray, P.B. and Garcia, J.R. Cambridge, Massachusetts: Harvard University Press, 2013.   Quarterly Review of Biology , 89 (3): 263-264. PDF

Puts, D.A.
,   Doll, L.M. , and   Hill, A.K.  (2014). Sexual selection on human voices. In:   Evolutionary Perspectives on Human Sexual Psychology and Behavior , Weekes-Shackelford, V. and Shackelford, T.K. (eds.). Springer, pp. 69-86.  PDF

Welling, L.L.M.
 and   Puts, D.A.  (2014). Female adaptations to ovulation. In:   Evolutionary Perspectives on Human Sexual Psychology and Behavior , Weekes-Shackelford, V. and Shackelford, T.K. (eds.). Springer, pp. 245-262. PDF

Wheatley, J.R.
, Apicella, C.L.,   Burriss, R.P. , Cárdenas, R.A., Bailey, D.H.,   Welling, L.L.M. , and   Puts, D.A.  (2014). Women’s faces and voices are cues to reproductive potential in industrial and forager societies.   Evolution and Human Behavior , 35(4): 264-271. PDF


Hill, A.K., Dawood, K. and Puts, D.A. (2013). Biological foundations of sexual orientation. In: Handbook of Psychology and Sexual Orientation, Patterson, C.J. and D'Augelli, A.R. (eds.). Oxford, UK: Oxford University Press, pp. 55-68. PDF

Hill, A.K., Hunt, J., Welling, L.L.M., Wheatley, J.R., Cárdenas, R.A., Rotella, M.A., Dawood, K., Shriver, M.D., and Puts, D.A. (2013). Quantifying the strength and form of sexual selection on men's traits. Evolution and Human Behavior, 34(5): 334-341. PDF

Kempe, V., Puts, D.A., and Cárdenas, R.A. (2013). Masculine men articulate less clearly. Human Nature, 24: 461-475. PDF

Puts, D.A., Bailey, D.H., Cárdenas, R.A., Burriss, R.P., Welling, L.L.M., Wheatley, J.R., and Dawood, K. (2013). Women's attractiveness changes with estradiol and progesterone across the ovulatory cycle. Hormones and Behavior, 63(1): 13-19. PDF

Puts, D.A. and Pope, L.E. (2013). Moderators, mates, and matchmakers: Effects of oral contraceptives on sexual desire may also depend on partners' behavior and the role of female choice. Archives of Sexual Behavior, 42(8). PDF

Welling, L.L.M.Persola, L.Wheatley, J.R., Cárdenas, R.A., and  Puts, D.A. (2013). Competition and men’s face preferences. Personality and Individual Differences, 54(3): 414-419. PDF

Welling, L.L.M.Singh, K.W.Puts, D.A., Jones, B.C., and Burriss, R.P. (2013). Self-reported sexual desire in homosexual men and women predicts preferences for sexually dimorphic facial cues. Archives of Sexual Behavior, 42(5): 785-791. PDF


Claes, P.M., Walters, M.D., Shriver, M.D.,  Puts, D.A., Gibson, G., Clement, J.G., Baynam, G.,Verbeke, G., Vandermeulen, D., and Suetens, P. (2012). Sexual dimorphism in multiple aspects of 3D facial symmetry and asymmetry defined by spatially dense geometric morphometrics. Journal of Anatomy, 221(2): 97-114. PDF

Deaner, R.O., Geary, D.C.,  Puts, D.A., Ham, S.A., Kruger, J., Fles, E., Winegard, B., and Grandis, T. (2012). A sex difference in the predisposition for physical competition: Males play sports much more than females even in the contemporary U.S.  PLoS ONE, 7(11): e49168. PDF

Puts, D.A., Apicella, C.L., and Cárdenas, R.A. (2012). Masculine voices signal men’s threat potential in forager and industrial societies. Proceedings of the Royal Society B: Biological Sciences, 279: 601-609. PDF

Puts, D.A. and Dawood, K. (2012). Is female orgasm a covert mate choice mechanism? In:  Oxford Handbook of Sexual Conflict in Humans, Shackelford, T.K. and Goetz, A. (eds.). Oxford University Press, pp. 184-201. PDF

Puts, D.A., Dawood, K., and  Welling, L.L.M. (2012). Why women have orgasms: An evolutionary analysis.  Archives of Sexual Behavior. PDF

Puts, D.A., Jones, B.C., and DeBruine, L.M. (2012). Sexual selection on human faces and voices.  Journal of Sex Research, 49 (2-3): 227-243. PDF

Puts, D.A.Welling, L.L.M.Burriss, R.P., and Dawood, K. (2012). Men’s masculinity and attractiveness predict their female partners’ reported orgasm frequency and timing.  Evolution and Human Behavior, 33: 1-9. PDF

Welling, L.L.M.Puts, D.A., Roberts, S.C., Little, A.C., and  Burriss, R.P. (2012). Hormonal contraceptive use and mate retention behavior in women and their male partners.  Hormones and Behavior, 61 (1): 114–120. PDF


Burriss, R.P., Roberts, S.C.,  Welling, L.L.M.Puts, D.A., and Little, A.C. (2011). Heterosexual romantic couples mate assortatively for facial symmetry, but not masculinity.  Personality and Social Psychology Bulletin, 37 (5): 601-613. PDF

Burriss, R.P.Welling, L.L.M., and  Puts, D.A. (2011). Mate-preference drives mate-choice: Men's self-rated masculinity predicts their female partner's preference for masculinity.  Personality and Individual Differences, 51 (8): 1023-1027. PDF

Burriss, R.P.Welling, L.L.M., and  Puts, D.A. (2011). Men's attractiveness predicts their preference for female facial femininity when judging for short-term, but not long-term, partners.  Personality and Individual Differences, 50 (5): 542-546. PDF

Hodges-Simeon, C.R., Gaulin, S.J.C., and  Puts, D.A. (2011). Voice correlates of mating success in men: Examining “contests” versus “mate choice” modes of sexual selection.  Archives of Sexual Behavior, 40 (3): 551-557. PDF

Puts, D.A.Barndt, J.L.Welling, L.L.M., Dawood, K., and  Burriss, R.P. (2011). Intrasexual competition among women: Vocal femininity affects perceptions of attractiveness and flirtatiousness. Personality and Individual Differences, 50(1): 111-115. PDF

Vukovic, J., Jones, B.C., Feinberg, D.R., DeBruine, L.M., Smith, F.G., Welling, L.L.M., and Little, A.C. (2011). Variation in perceptions of physical dominance and trustworthiness predicts individual differences in the effect of relationship context on women's preferences for masculine pitch in men's voices.  British Journal of Psychology, 102 (1): 37-48. PDF

Welling, L.L.M.Burriss, R.P., and  Puts, D.A. (2011). Mate retention behavior modulates men’s preferences for self-resemblance in infant faces.  Evolution and Human Behavior, 32 (2): 118-126. PDF


Bestelmeyer, P.E.G., Jones, B.C., DeBruine, L.M., Little, A.C., and  Welling, L.L.M. (2010). Face aftereffects suggest interdependent processing of expression and sex and of expression and race.  Visual Cognition, 18(2): 255-274. PDF

Burriss, R.P. (2010). The effect of veneers on cosmetic improvement: comments on Nalbandian and Millar, 2009.  British Dental Journal, 208(2): 47. PDF

DeBruine, L.M., Jones, B.C., Crawford, J.R.,  Welling, L.L.M., and Little, A.C. (2010). The health of a nation predicts their mate preferences: Cross-cultural variation in women's preferences for masculinized male faces.  Proceedings of the Royal Society B: Biological Sciences, 277(1692): 2405-2410. PDF

DeBruine, L.M.,  Welling, L.L.M., Jones, B.C., and Little, A.C. (2010). Opposite effects of visual versus imagined presentation of faces on subsequent sex perception.  Visual Cognition, 18(6): 816-828. PDF

Grimbos, T., Dawood, K.,  Burriss, R.P., Zucker, K.J., and  Puts, D.A. (2010). Sexual orientation and the second to fourth finger length ratio: A meta-analysis in men and women.  Behavioral Neuroscience, 124(2): 278-287. PDF

Hodges-Simeon, C.R., Gaulin, S.J.C., and  Puts, D.A. (2010). Different vocal parameters predict perceptions of dominance and attractiveness.  Human Nature, 21(4): 406-427. PDF

Jones, B.C., DeBruine, L.M., Main, J.C., Little, A.C.,  Welling, L.L.M., Feinberg, D.R., and Tiddeman, B.P. (2010). Facial cues of dominance modulate the short-term gaze-cuing effect in human observers.  Proceedings of the Royal Society B: Biological Sciences, 277: 617-624. PDF

Jones, B.C., Main, J.C., DeBruine, L.M., Little, A.C., and  Welling, L.L.M. (2010). Reading the look of love: Sexually dimorphic cues in opposite-sex faces influence gaze categorization.  Psychological Science, 21(6): 796-798. PDF

Puts, D.A. (2010). Beauty and the beast: Mechanisms of sexual selection in humans.  Evolution and Human Behavior, 31(3): 157-175. PDF

Puts, D.A. (2010). The psychologic gambit declined—a review of “Endocrinology of Social Relationships.”  Evolution and Human Behavior, 31(4): 306-308. PDF

Puts, D.A., Cárdenas, R.A., Bailey, D.H.,  Burriss, R.P., Jordan, C.L., and Breedlove, S.M. (2010). Salivary testosterone does not predict mental rotation performance in men or women.  Hormones and Behavior, 58(2): 282-289. PDF

Vukovic, J., Jones, B.C., DeBruine, L.M., Feinberg, D.R., Smith, F.G., Little, A.C.,  Welling, L.L.M., and Main, J.C. Women's own voice pitch predicts their preferences for masculinity in men's voices.  Behavioral Ecology, 21(4): 767-772. PDF

Wolff, S.E., and  Puts, D.A. (2010). Vocal masculinity is a robust dominance signal in men.  Behavioral Ecology and Sociobiology, 64(10): 1673-1683. PDF


Burriss, R.P. (2009). Esthetic effect of orthodontic appliances: comments on Berto et al., 2009.  American Journal of Orthodontics & Dentofacial Orthopedics, 136(3): 305-306. PDF

Burriss, R.P. (2009). Symmetry is sexy: Reply to Hodgson’s ‘Symmetry and Humans’.  Antiquity, 83(322): 1170-1175. PDF

Burriss, R.P., Rowland, H.M., and Little, A.C. (2009). Facial scarring enhances men’s attractiveness for short-term relationships.  Personality and Individual Differences, 46: 213-217. PDF

Puts, D.A.  (2009).  The Evolution of Human Sexuality: An Anthropological Perspective (2nd ed) . Dubuque, Iowa: Kendall/Hunt.  View at

Roberts, S.C., Saxton, T.K., Murray, A.K.,  Burriss, R.P., Rowland, H.M., and Little, A.C. (2009). Static and dynamic facial images cue similar attractiveness judgements.  Ethology, 115: 588-595. PDF

Saxton, T.K.,  Burriss, R.P., Murray, A.K., Rowland, H.M., and Roberts, S.C. (2009). Face, body and speech cues independently predict judgments of attractiveness.  Journal of Evolutionary Psychology, 7: 23-35. PDF

Smith, F.G., DeBruine, L.M., Jones, B.C., Krupp, D.B.,  Welling, L.L.M., and Conway, C.A. (2009). Attractiveness qualifies the effect of observation on trusting behavior in an economic game.  Evolution and Human Behavior, 30(6): 393-397. PDF

Smith, F.G., Jones, B.C., Little, A.C., DeBruine, L.M.,  Welling, L.L.M., Vukovic, J., and Conway, C.A. (2009). Hormonal contraceptive use and perceptions of trust modulate the effect of relationship context on women's preferences for sexual dimorphism in male face shape. Journal of Evolutionary Psychology, 7(3): 195-210. PDF

Wolff, S.E., and  Puts, D.A. (2009). Book review:  Sex Differences: Summarizing More Than a Century of Scientific Research, by L. Ellis, S. Hershberger, E. Field, S. Wesinger, S. Pellis, D. Geary, C. Palmer, K. Hoyenga, A. Hetsroni, and K. Karadi. New York: Psychology Press, 2008.  Archives of Sexual Behavior, 38(6): 1070. PDF


Dawood, K. and  Puts, D.A.  (2008). Genetics and the development of human sexual orientation. In:  LGBTQ America Today , J.C. Hawley (ed.). Westport, Connecticut: Greenwood Press, pp. 469-477. PDF

Martin, J.,  Puts, D.A., and Breedlove, S.M. (2008). Hand asymmetry in heterosexual and homosexual men and women: Relationship to 2D:4D digit ratios and other sexually dimorphic anatomical traits. Archives of Sexual Behavior, 37(1): 119-132. PDF

Puts, D.A., McDaniel, M.A., Jordan, C.L., and Breedlove, S.M. (2008). Spatial ability and prenatal androgens: Meta-analyses of congenital adrenal hyperplasia and digit ratio (2D:4D) studies.  Archives of Sexual Behavior, 37(1): 100-111. PDF

Zuloaga, D.G.,  Puts, D.A., Jordan, C.L., and Breedlove, S.M. (2008). The role of androgen receptors in the masculinization of brain and behavior: What we've learned from the testicular feminization mutation. Hormones and Behavior, 53(5): 613-626. PDF


Puts, D.A.  (2007). Arousing imaginations: Review of  Handbook of the Evolution of Human Sexuality , M.R. Kauth (ed.). Binghamton, NY: Haworth Press, 2006.  Evolutionary Psychology , 5(4): 778-785.  PDF

Puts, D.A. (2007). Of bugs and boojums: Female orgasm as a facultative adaptation.  Archives of Sexual Behavior, 36(3): 337-339. PDF

Puts, D.A. , Gaulin, S.J.C., and Breedlove, S.M. (2007). Sex differences in spatial ability: Evolution, hormones, and the brain. In: Evolutionary Cognitive Neuroscience , S.M. Platek, J.P. Keenan, and T.K. Shackelford (eds.). Cambridge, Massachusetts: MIT Press, pp. 329-379.  PDF

Puts, D.A., Hodges, C.R., Cárdenas, R.A., and Gaulin, S.J.C. (2007). Men’s voices as dominance signals: Vocal fundamental and formant frequencies influence dominance attributions among men.  Evolution and Human Behavior, 28(5): 340-344. PDF


Puts, D.A. (2006). And hast thou slain the Jabberwock? Response to Wallen.  Archives of Sexual Behavior, 35(6): 637-639. PDF

Puts, D.A. (2006). Review of  The Case of Female Orgasm: Bias in the Science of Evolution, by E.A. Lloyd. Cambridge, Massachusetts: Harvard University Press, 2005.  Archives of Sexual Behavior, 35(1): 103-108. PDF

Puts, D.A. (2006). Cyclic variation in women’s preferences for masculine traits: Potential hormonal causes.  Human Nature, 17(1): 114-127. PDF

Puts, D.A. and Dawood, K. (2006). The evolution of female orgasm: Adaptation or byproduct?  Twin Research and Human Genetics, 9(3): 467-472. PDF

Puts, D.A., Gaulin, S.J.C., and Verdolini, K. (2006). Dominance and the evolution of sexual dimorphism in human voice pitch.  Evolution and Human Behavior, 27(4): 283-296. PDF

Puts, D.A., Jordan, C.L., and Breedlove, S.M. (2006). Defending the brain from estrogen.  Nature Neuroscience, 9(2): 155-156. PDF

Puts, D.A., Jordan, C.L., and Breedlove, S.M. (2006). O brother, where art thou? The fraternal birth-order effect on male sexual orientation.  Proceedings of the National Academy of Sciences of the USA, 103(28): 10531-10532. PDF

2005 and earlier

Puts, D.A. (2005). Mating context and menstrual phase affect women’s preferences for male voice pitch.  Evolution and Human Behavior, 26(5): 388-397. PDF

Puts, D.A., Gaulin, S.J.C., Sporter, R.J., and McBurney, D.H. (2004). Sex hormones and finger length: What does 2D:4D indicate?  Evolution and Human Behavior, 25(3): 182-199. PDF

Puts, D.A.  (2003).  Human Sexuality: A Holistic Approach . Dubuque, Iowa: Kendall/Hunt.

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